Beskrivelse: Beskrivelse: Danish Logo

Beskrivelse: Beskrivelse: indeksHome .Beskrivelse: Beskrivelse: indeks. Menu

Genus Rhododendron

Inspireret af Goetsch et al. 2005  

Goetsch et al.2005 classification used mostly non-coding sequencies in RPB2-I. Exon areas is cut out before use (B. Hall 2007). 
ITS do not have important sequences. RPB2-I and ITS have more gaps and therefore difficult to work with. No gaps in matK coding area. 
RPB2-I sequence exon23-24 gave a lot of "noise" using some programes, mostly from parts of exon 23 and 24 if exons is not cut out.
Goetsch et al. data are very informative and the largest to day and the "noise" give only minor problems?
At the moment the Menziesia group is in this Genus. R. camtchaticum is a matter of feelings  

 


Example of a phylogenetic tree using ITS from 1998:
Phylogenetic tree maked by using DNA from the ITS area

H. Eiberg.  Rhododendronfamiliens stamtræ.  In: Rhododendron i Danmark i 25 år.Nielsen BL, Hansen SM. (1999). p.186-206. RhodoNyt 1997(1): p 14-18.  www.rhododendron.dk/taxo.htm

 

 -yellow highlights  is something I think about   1/4-2007 H. Eiberg  -  Mange har nu fået et gruppenavn (sec./subsec.) i Flora of China      ** komentarer.

 
 
 Flora of China’s klassifikation er delvist bygget på DNA analyser.
 
Classification after Flora of China  2010        Arter alfabetisk: http://www.efloras.org/browse.aspx?flora_id=2&start_taxon_id=128386&page=1  
Flora of China's classification is partly built on DNA analysis.

 
 
 
 
 
 
 
New species, variants, forms  are added after:  Pocket Guide to Rhododendron based on the descriptions by H.H. Davidian. By J.F.J. Mc Quire and M.L.A. Robinson. Kew 2009 
 
 <-var = .* As species in Flora of China
   

 

R. chamberlainii  (Diplarche multiflora)       Perhaps closer to rhododendron than R. camtschaticum? (HE)

 

 

(Subgenus Menziesia (HE) (If  Therorhodion is in this genus, then Menziesia is (HE, 1997) - also after  DNA studies of several DNA regions)
 
            R. goyozanense (Menziesia goyozanensis)
                       R. kamadae     (Menzisia kamadae)
                       R. katsumatae  (Menziesia katsumatae)
            R. kroniae
                       R. multiflorum  (Menziesia ciliicalyx var. multiflora)
                       R. multiflorum var. purpureum  (Menziesia multiflora var. purpurea)
                       R. multiflorum forma brevicalyx
                       R. multiflorum forma bicolor (M. multiflora var. bicolor, M. lasiophylla)
                       R. multiflorum forma longicalyx (M. multiflora var. longicalyx)
                       R. menziesii   (Menzisia ferruginea)
                       R. menziesii ssp. glabellum  (Menzisia ferruginea ssp. glabella)
                       R. pentandrum  (Menzisia pentandra)
                       R. pilosum     (Menzisia pilosa)
                       R. sophistarum
                       R. yakushimense  (Menziesia lasiophylla yakushimensis)
 
 
Eiberg, Hans (1997) Rhododendronfamiliens stamtræ 2. Rhodo-Nyt 1: 14-18
Eiberg, Hans (1999) Rhododendronslægtens stamtræ. in Rhododendron i Danmark i 25 år page 186-199. Copenhagen.
 ----------
 
Craven, L.A. (2011). Diplarche and Menziesia transferred to Rhododendron (Ericaceae). Blumea 56: 33-35.
 
Abstract:  The genera Diplarche and Menziesia have been shown to be nested in Rhododendron based upon cladistic analyses of DNA data. 
Morphologically, the differences between Diplarche and Menziesia are not so great that their inclusion in Rhododendron is 
inappropriate and the species of these two genera are formally transferred to Rhododendron. The following new names are proposed: 
Rhododendron benhallii, R. chamberlainii, R. goyozanense, R. × kamatae, R. katsumatae, R. kroniae, R. menziesii, 
R. menziesii subsp. glabellum, R. multiflorum, R. multiflorum forma bicolor, R. multiflorum forma brevicalyx, 
R. multiflorum var. purpureum, R. pentandrum, R. pilosum, R. sophistarum, R. yakushimense.
 
  ---------

Subgenus Azaleastrum       (clade C*)

                     Section Sciadorhodion

 

               Subsection Candidastrum(HE)
                       R. albiflorum
 
               
 
               Subsection Schlippenbachia (J. Batta 1980)HE
                       R. albrechtii                  
                       R. pentaphyllum
                       R. quinquefolium
                       R. schlippenbachii
 
               Subsection Vaseya(HE)
                       R. vaseyi
 
               Subsection Nipponica (J. Batta 1980)HE ßà Tsutsusi
                       R. nipponicum  

 

                     Section Tsutsusi

               Subsection Brachycalyces (Spethmann 1987)HE
                   Farrerae alliance   (sec. Brachycalyx*)
                       R. chilanshanense*     
                       R. farrerae
                       R. huadingense*
                       R. mariesii
 
                   Reticulatum alliance       
                       R. reticulatum
                       R. decandrum
                       R. dilatatum
                       R. hidakanum
                       R. kiyosumense
                       R. mayebarae
                       R. nudipes
                               ssp. nudipes
                               ssp. niphophilum var. lagopus
                       R. viscistylum
                       R. wadanum
                  Weyrichii alliance  
                       R. amagianum
                       R. sanctum

                       R. weyrichii

 
 
               Subsection Mumeazalea (M.N.Philipson & Philipson 1982)(HE)
 
                       R. semibarbatum
               
               Subsection ovata(HE) (now sec. Azaleastrum in Flora of China)  
                       R. bachii*
                       R. hongkongense
                       R. leptothrium
                       R. medoense*
                       R. mitriforme*
                                              var. mitriforme* 
                                              var. setaceum*
                       R. ovatum
                       R. vialii
 

 

               Subsection Tsutsusi(HE)  In Flora of China*
 
 
Author(s): Jin XiaoFeng; Ding BingYang; Jin ShuiHu; Zhang YueJiao; Fu ChengXin
Title: Revision of some problematic taxa of Rhododendron sect. Tsutsusi (Ericaceae) from China.
Source: Annales Botanici Fennici 44 (1) : 18-24 2007
Abstract: Morphological variation within and among populations of some closely 
related taxa of Rhododendron sect. Tsutsusi from China are revised based on 
literature, specimen examination and field work. Rhododendron chunii, R. guizhongense,
R. viscidum, R. subenerve, R. minutiflorum, R. viscigemmatum, and R. polyraphidodeum
var. montanum are reduced to synonymys of R. fuchsiifolium. Rhododendron myrsinifolium
is recognized as a distinct species, differing from R. fuchsiifolium in having smaller
leaves, branchlets covered with compressed strigose hairs, and a corolla tube
without glands on outer surface. The geographic distributions of R. fuchsiifolium
and R. myrsinifolium are also provided.
 
 
                       R. adenanthum*
                       R. atrovirens
                       R. boninense
                       R. breviperulatum
                       R. chunii*
                       R. crassimedium*
                       R. crassistylum*
                       R. cretaceum*
                       R. eriocarpum
                       R. flosculum*
                       R. flumineum*   --> R. mariae ssp. flumineum
                       R. fuchsiifolium*
                       R. fuscipilum*
                       R. guizhongense*
                       R. hainanense*
                       R. hejiangense*
                       R. hunanense*
                       R. hypoblematosum*
                       R. indicum
                       R. jinpingense*
                       R. jinxiuense*
                       R. kaempferi
                       R. kanehirae
                       R. kiusianum           var. kiusianum
                                              var. sataense
                       R. kwangtungense*
                       R. lasiostylum*
                       R. lilacinum     (new 2010)
                       R. linguiense*
                       R. longifalcatum*
                       R. longiperulatum*
                       R. loniceriflorum*
                       R. macrosepalum
                       R. malipoense*    --> R. mariae subsp. flumineum
                       R. microphyton    --> R. mariae ssp. microphyton
                       R. minutiflorum*
                       R. mucronatum          var. ripense
                                              var. mucronatum
                       R. myrsinifolium*
                       R. nakaharae
                       R. nanpingense*
                       R. noriakianum
                       R. octandrum*
                       R. oldhamii **
                       R. polyraphidoideum*
                       R. pulchroides*
                       R. qianyangense*
                       R. rhodanthum*
                       R. rhuyuenense*
                       R. rufohirtum*
                       R. rubropilosum
                       R. saxicolum
                       R. saxatile*
                       R. scabrum
                       R. seniavinii*
                       R. serpyllifolium
                       R. simsii
                       R. spadiceum*
                       R. sparcifolium*
                       R. strigosum*
                       R. subflumineum*
                       R. subnerve*
                       R. subserinum*
                       R. subsessile
                       R. taipaoense*
                       R. taiwanalpinum*
                       R. tashiroi
                       R. temilaminare*
                       R. tenue*
                       R. tingwuense*
                       R. tosaense
                       R. tschonoskii
                       R. tsoi*
                       R. tsusiophyllum
                       R. unciferum*
                       R. viscidum*
                       R. viscigemmatum*
                       R. yangmingshanense*
                       R. yaoshanicum*
                       R. yedoense var. poukhanense
                       
Subgenus Choniastrum
 
                       R. championiae
                       R. hancockii
                                              var. hancockii
                                              var. longisepalum*
                       R. latoucheae
                       R. moulmainense
                                                                     ellipticum
                       R  stamineum
                                              var. gaozhaiense*
                                              var. lasiocarpum*
                                              var. stamineum*
                    --add*
 
                       R. cavaleriei*
                       R. dayaoshanense*
                       R. feddei*
                       R. henryi*
                                              var. dunnii*
                                              var. henryi*
                       R. huguangense*
                       R. longilobum*
                       R. mackenzianum*
                       R. subestipitatum*
                       R. taishuense*
                       R. truncatovarium*
                       R. tutcherae*
                                              var. glabrifolium*
                                              var. gymnocarpum*
                                              var. tutcherae*
                       R. vaniotii*
                       R. westlandii*
 

Subgenus Hymenanthes

                     Section Pentanthera. Could be a subgenus (HE)   In: Flora of China* it is still a subgenus

 

               Subsection Canadensia (J. Batta 1980, Spethmann 1987) HE                     
                       R. canadense
 
               Subsection Lutea (J. Batta 1980, Spethmann 1987) (HE)
                       R. alabamense
                       R. arborescens
                       R. atlanticum
                       R. austrinum
                       R. calendulaceum
                       R. canescens
                       R. colemanii
                       R. cumberlandense
                       R. eastmanii
                       R. flammeum
                       R. luteum
                       R. occidentale
                       R. periclymenoides
                                              var. glandiferum
                       R. prinophyllum
                       R. prunifolium
                       R. viscosum
                                              var. glaucum
                                                             oblongifolium
                                                             serrulatum
                       
               Subsection Sinensia (K.A. Kron 1993) HE
 
                       R. molle       ssp. molle
                                      ssp. japonicum
         

                     Section Ponticum

               Subsection Arborea (RSBG: Rhododendron Species 2011, 89-100)
 
                       R. arboreum    
                                      ssp. arboreum
                                      ssp. cinnamomeum 
                                              var. cinnamomeum
                                              var. roseum
 
                       R. delavayi    
                                              var. adenostylum (new 2009)
                                              var. delavayi
                                              var. peramoenum
                                              var. pilostylum*
                                              var. albotomentosum
                                              R. delavayi aff.
                       R. nilagiricum
                       R. zeylanicum
                       R. lanigerum
                       R. niveum
 
               Subsection Argyrophylla
 
                       R. adenopodum
                       R. argyrophyllum    
                                      ssp. argyrophyllum
                                      ssp. hypoglaucum à R. hypoglaucum*
                                      ssp. omeiense
                                      ssp. nankingense
                                              
                       R. brevipetiolatum*
                       R. coryanum
                       R. denudatum
                                                 var. glabriovarium
                       R. ebianense*
                       R. eriobotryoides (new 2009)
                       R. fangchengense*
                       R. farinosum
                       R. floribundum
                       R. formosanum
                       R. haofui
                       R. hunnewellianum
                                              var. hunnevellianum*
                                              var. rockii*
                       R. insigne
                                              var. hejiangense*
                                              var. insigne*
 
                       R. longipes            var. chienianum
                                              var. longipes
                       R. oblancifolium*
                       R. pingianum
                       R. ririei
                       R. shimenense*
                       R. simiarum
                                              var. simiarum*
                                              var. versicolor*
                       R. thayerianum
 
               Subsection Auriculata
 
                       R. auriculatum
                       R. chihsinianum*
 
               Subsection Barbata
 
                       R. argipeplum
                       R. barbatum
                       R. erosum
                       R. exasperatum
                       R. succothii 
 
               Subsection Campanulata
 
                       R. campanulatum 
                                      ssp. campanulatum
                                      ssp. aeruginosum
                       R. gannanense*
                       R. wallichii
                                                     heftii
 
               Subsection Campylocarpa
 
                       R. callimorphum        var. callimorphum  
                                                          cyclium
                                              var. myiagrum
                       R. campylocarpum   
                                      ssp. campylocarpum
                                      ssp. caloxanthum
                                                     telopeum 
                                                     panteumorphum
                       R. henanense*
                                      ssp. henanense*
                                      ssp. lingbaoense*
                       R. longicalyx*
                       R. souliei
                       R. wardii              var. wardii
                                                     litiense
                                              var. puralbum
               Subsection Falconera
 
                       R. arizelum
                       R. basilicum
                       R. coriaceum
                       R. falconeri
                                      ssp. eximium
                                      ssp. falconeri
                       R. galactinum
                       R. heatheriae
                       R. hodgsonii
                       R. hodgsonii  aff.                            
                       R. preptum
                       R. rex         ssp. rex
                                      ssp. fictolacteum   
                                              var. miniforme
                                      ssp. gratum
                       R. rothschildii
                       R. semnoides
                       R. sinofalconeri
 
               Subsection Fortunea
 
                       R. asterochnoum
                                              var. brevipedicellatum*
                       R. calophytum  
                                              var. calophytum
                                              var. jinfuense*
                                              var. openshawianum
                                              var. pauciflorum*
                       R. davidii
                       R. decorum
                                      ssp. decorum
                                      ssp. diaprepes
                       R. faithiae*
                       R. fortunei
                                      ssp. fortunei
                                      ssp. discolor
                                                             f. cordatum
                       R. gingongshanicum ?
                       R. glanduliferum
                       R. gonggashanense*
                       R. griffithianum
                       R. guihainianum*
                       R. hemsleyanum
                                              var. chengianum*
                                              var. hemsleyanum*
                       R. huanum*
                      R. huangpingense (new 2010)
                       R. huianum /huanum
                       R. jinchangense
                       R. jingangshanicum*
                       R. jiulongshanense (new 2009)
                       R. magniflorum*
                       R. maoerense*
                       R. miyiense*
                       R. orbiculare
                                      ssp. oblongum*
                                      ssp. orbiculare
                                      ssp. cardiobasis
                       R. oreodoxa
                                              var. adenostylosum*
                                              var. oreodoxa
                                              var. fargesii 
                                              var. shensiense
                       R. platypodum 
                       R. praeteritum
                       R. praevernum
                       R. qiaojiaense
                       R. serotinum
                       R. sutchuenense                
                       R. vernicosum
                                                             f. araliiforme
                                                             f. euanthum
                                                             f. rhantum
                                                             f. sheltonae
                       R. verruciferum*
                       R. wolongense*
                       R. xiaoxodongense*
                       R. yuefengense
                       
               Subsection Fulgensia 
 
                       R. fulgens
                       R. miniatum   *(moved)
                       R. sherriffii *(moved)
 
               Subsection Fulva
 
                       R. fulvum      
                                      ssp. fulvum
                                      ssp. fulvoides
                       R. uvarifolium
                                              var. griseum
                                              var. uvarifolium
 
               Subsection Glischra            
 
                       R. adenosum
                       R. crinigerum
                                              var. crinigerum
                                              var. euadenium
                       R. diphrocalyx ->X*
                       R. glischroides
                                              var. arachnoideum
                       R. glischrum
                                      ssp. glischrum
                                      ssp. rude
                       R. habrotrichum
                       R. recurvoides
                       R. spilotum
                       R. vesiculiferum
 
               Subsection Grandia
 
                       R. balangense  or Taliensa
                       R. grande
                       R. kesangiae           var. kesangiae
                                              var. album
                       R. macabeanum 
                       R. magnificum
                       R. montroseanum
                       R. oreogenum*
                       R. praestans
                       R. protistum
                                              var. giganteum
                                              var. protistum*
                       R. pudorosum
                       R. sidereum
                       R. sinogrande
                       R. suoilenhense
                       R. watsonii
        
               Subsection Griersoniana  (monotypic)
 
                       R. griersonianum
                        
               Subsection Irrorata
 
                       R. aberconwayi
                       R. x agastum*
                                              var. agastum*
                                              var. pennivenium*
                       R. annae
                                      ssp. annae
                                      ssp. laxiflorum
                       R. anthosphaerum
                                                             eritimum
                       R. araiophyllum
                                              var. araiophyllum*
                                              var. lapidosum*
                       R. brevinerve*
                       R. dimiatrum*
                       R. gongshanense*
                       R. guizhouense*
                       R. irroratum   ssp. irroratum
                                      ssp. pogonostylum
                                      ssp. yiliangense
                       R. jinboense (new 2009)
                       R. kendrickii
                       R. langbianense
                       R. laojunshanense*
                       R. leptopeplum*
                       R. lukiangense
                       R. mengtszense*
                       R. ningyuenense
                       R. papillatum*
                       R. pingbianense*
                       R. ramsdenianum
                       R. sikangense* (from Maculifera*)
                       R. spanotrichum*
                       R. tanastylum          var. tanastylum
                                              var. pennivenium  
                                                                            x agastum*
                                              var. lingzhiense*
                       R. wrayi
                       R. ziyuanense
 
               Subsection Lanata
 
                       R. flinckii
                       R. lanatoides  
                       R. lanatum
                       R. luciferum  (Hobbiense)  
                       R. tsariense
                                              var. trimoense          
               Subsection Maculifera
 
                       R. anwheiense
                       R. leishanicum
                       R. longesquamatum
                       R. maculiferum
                       R. morii
                       R. ochraceum
                                              var. brevicarpum*
                                              var. ochraceum*
                       R. oligocarpum*
                       R. pachyphyllum*
                       R. pachysanthum
                       R. pachytrichum
                                              var. pachytrichum*
                                              var. tenuistylum*
                       R. pilostylum*
                       R. polytrichum*
                       R. pseudochrysanthum
                       R. sikangense          var. sikangense
                                              var. exquisitum
                       R. strigillosum
                                              var. strigillosum*
                                              var. monosematum*
                       R. subroseum
                       R. ziyuanense* 
 
               Subsection Neriiflora
 
               Haematodes alliance    
                       R. beanianum
                       R. catacosmum
                       R. chionanthum*
                       R. coelicum
                       R. haematodes
                                      ssp. chaetomallum
                                      ssp. haematodes
                                                                     x hemigynum
                       R. miniatum
                       R. piercei
                       R. pocophorum          var. pocophorum
                                              var. hemidartum
               Mallotum alliance      
                       R. mallotum
               Sanguineum alliance
                       R. aperantum
                       R. citriniflorum       var. citriniflorum
                                              var. horaeum 
                                                     var. rubens
                       R. dichroanthum
                                      ssp. apodectum
                                      ssp. dichroanthum
                                      ssp. scyphocalyx
                                      ssp. septentrionale
                       R. erastum*
                       R. eudoxum             var. eudoxum
                                              var. brunneifolium
                                              var. mesopolium
                       R. microgynum
                       R. parmulatum
                       R. sanguineum
                                      ssp. didymum  ->var.*
                                      ssp. sanguineum  ->var*
                                              var. sanguineum
                                              var. haemaleum
                                              var. himertum*
                                              var. cloiophorum
                                              var. didymoides
                       R. temenium
                                              var. dealbatum
                                              var. gilvum
                                              var. temenium
               Forrestii alliance
                       R. chamaethomsonii     
                                              var. chamaedoron
                                              var. chamaethauma
                                              var. chamaethomsonii
                       R. forrestii   ssp. forrestii
                                      ssp. papillatum
                       R. trilectorum*
               Neriiflorum alliance   
                       R. albertsenianum
                       R. bijiangense*
                       R. euchroum
                       R. floccigerum
                       R. neriiflorum
                                              var. appropinquans*
                                      ssp. agetum ->var.*
                                      ssp. neriiflorum ->var.*
                                      ssp. phaedropum ->var.
                       R. sperabile           var. sperabile
                                              var. weihsiense
                       R. sperabiloides
 
               Subsection Parishia
 
                       R. agapetum
                       R. elliottii 
                       R. facetum
                       R. flavoflorum*
                       R. huidongense*
                       R. kyawii
                       R. schistocalyx*
                       R. urophyllum*
                       R. venator** from Venata*
 
               Subsection Pontica
 
                       R. aureum
                                              var. aureum
                                              var. hypopytis
                       R. brachycarpum
                                      ssp. brachycarpum
                                              var. tigerstedtii
                                      ssp. fauriei
                       R. catawbiense
                       R. caucasicum
                       R. degronianum
                                      ssp. degronianum
                                      ssp. heptamerum 
                                              var. heptamerum
                                              var. hondoense
                                              var. kyomaruense
                                      ssp. yakushimanum
                       R. hyperythrum
                       R. macrophyllum
                       R. makinoi
                       R. maximum
                       R. ponticum
                       R. smirnowii
                       R. ungernii
 
               Subsection Selensia
                       
                       R. bainbridgeanum
                       R. calvescens*
                                              var. calvescens*
                                              var. duseimatum*
                       R. dasycladoides*
                                                                     R. X erythrocalyx
                       R. esetulosum
                       R. hirtipes
                       R. martinianum
                       R. selense
                                      ssp. dasycladum
                                      ssp. selense   
                                              var. pagophilum
                                      ssp. jucundum
                                              var. probum
                                      ssp. setiferum -> R. setiferum*
                       R. sertiferum*
                 
               Subsection Taliensia
 
                       R. adenogynum
                       R. aganniphum
                                              var. aganniphum
                                                                     glaucopeplum
                                              var. flavorufum 
                                              var. schizopeplum*     doshangense
                       R. alutaceum
                                              var. alutaceum
                                              var. iodes
                                              var. russotinctum
                       R. balangense* or Grandia
                       R. balfourianum
                                              var. aganniphoides*
                       R. barkamense*
                       R. x bathyphyllum*
                       R. bhutanense
                       R. bureavii
                       R. bureavioides
                       R. clementinae
                                      ssp. aureodorsale*
                       R. circinnatum
                       R. codonanthum*
                       R. coeloneurum
                       R. comisteum*
                       R. dachengense*
                                              var. dachengense*
                                              var. scopulum*
                       R. danbaense*
                       R. detersile
                       R. x detonsum
                       R. dumicola
                       R. elegantulum
                       R. faberi
                                      ssp. prattii*
                       R. heizhugouense*
                       R. lulangense*
                       R. mimetes             var. mimetes
                                              var. simulans
                       R. montiganum*
                       R. nakotiltum*
                       R. nigroglandulosum
                                                                            x paradoxum
                       R. phaeochrysum 
                                              var. agglutinatum
                                              var. levistratum     dryophyllum
                                              var. phaerochrysum
                       R. pomense*
                       R. potaninii*
                       R. prattii àfaberi ssp. prattii*
                       R. principis
                       R. pronum
                       R. proteoides
                       R. przewalskii
                                      ssp. chrysophyllum*
                                      ssp. dabanshanense
                                      ssp. huzhuense*
                                      ssp. przewalskii
                                      ssp. yushuense*
                       R. pubicostatum*
                       R. pudiense (new 2009)
                       R. pugeense*
                       R. punctifolium*
                       R. purdomii
                       R. roxieanum
                                              var. cucullatum
                                                                     deep rose
                                              var. oreonastes
                                              var. roxieanum
                                                             ? recurvum
                                                                            x parvum
                       R. roxieoides*
                       R. rufum
                                                                     weldianum
                       R. shanii*
                       R. sphaeroblastum      var. wumengense
                       R. taliense            
                                              aff.
                       R. torquescens*
                       R. traillianum
                                              var. dictyotum
                                              var. traillianum
                       R. trichogynum*
                       R. wasonii
                                              var. wasonii
                                              var. wenchuanense
                       R. wasonii aff. 
                       R. wiltonii
                       R. zhongdianense*
 
                       R. x inopinum          
                       R. x paradoxum
               Lacteum alliance       
                       R. beesianum
                       R. dignabile
                       R. lacteum
                       R. nakotiltum*
                       R. wightii
 
               Subsection Thomsonia
 
                       R. bonvalotii*
                       R. cerasinum
                       R. cyanocarpum
                       R. eclecteum
                                              var. bellatulum
                       R. eurysiphon*
                       R. faucium     
                       R. hookeri
                       R. hylaeum
                       R. meddianum
                                              var. atrokermesinum
                                              var. meddianum
                       R. megalanthum*
                       R. populare*
                       R. ramipilosum*
                       R. sherriffii ->Fulgensia*
                       R. stewartianum
                       R. subansiriense
                       R. thomsonii   ssp. thomsonii
                                      ssp. lopsangianum
                       R. viscidifolium
 
               Subsection Venatora  (monotypic) -> Parishia*
 
                       R. venator  -> Parishia*
                       
               Subsection Williamsiana  
 
                       R. williamsianum
                       (R. leishanicum*)  -> Maculifera
 

Subgenus Rhododendron

        Section Pogonanthum
 
                       R. anthopogon
                                      ssp. anthopogon  ->R. anthopogon*
                                      ssp. hypenanthum ->R. hypenanthum*  
                       R. anthopogonoides
                       R. atropunicum*
                       R. bellissimum*
                       R. cephalanthum 
                                      ssp. cephalanthum
                                                                     nmaiense
                                      ssp. platyphyllum ->R. platyphyllum*
                       R. collettianum
                       R. fragrans (adamsii)
                       R. heteroclitum*
                       R. hoi
                       R. kongboense
                       R. laudandum           var. laudandum
                                              var. temoense
                       R. luhuoense*
                       R. mainlingense*
                       R. nyingchiense*
                    R. pogonophyllum
                       R. primuliflorum
                                              var. cephalantoides*
                                              var. lepidanthum*
                                              var. primuliflorum*
                                                                     clivicola
                       R. radendum*
                       R. rufescens
                       R. sargentianum
                       R. trichostomum
                                                     f. hedyosmum
                                                     f. radinum
                                                                     ledoides
                       R. tubulosum*
                       
        Section Rhododendron
 
               Subsection Afghanica  (monotypic)
 
                       R. afghanicum
 
               Subsection Baileya  (monotypic)
 
                       R. baileyi
 
               Subsection Boothia
 
                       R. boothii
                                                             mishmiense
                       R. chrysodoron
                       R. dekatanum
                       R. leptocarpum
                       R. leucaspis
                       R. megeratum
                       R. nanjianense
                       R. sulfureum
 
               Subsection Camelliiflora  (monotypic)
 
                       R. camelliiflorum
 
               Subsection Campylogyna  (monotypic)
 
                       R. campylogynum
                                                             celsum 
                                                             charopoeum 
                                                             myrtilloides 
                                                             cremastum 
                                                                     Leucanthum
 
               Subsection Caroliniana  (monotypic)
 
                       R. minus
                                              var. chapmanii
                                              var. minus
               Subsection Cinnabarina
 
                       R. cinnabarinum 
                                      ssp. cinnabarinum
                                              var. aestivale
                                              var. blandfordiiflorum
                                              var. cinnabarinum
                                                             breviforme
                                              var. purpurellum
                                              var. roylei            
                                      ssp. tamaense
                                      ssp. xanthocodon   
                                              var. concatenans
                       R. igenum*
                       R. keysii
                       R. lateriflorum*
                       R. tenuifolium*
 
               Subsection Edgeworthia
 
                       R. edgeworthii
                       R. pendulum
                       R. seinghkuense
 
               Subsection Fragariflora  (monotypic)
 
                       R. fragariiflorum
 
               Subsection Genestieriana  (monotypic)
 
                       R. genestierianum
 
               Subsection Glauca  
 
                       R. brachyanthum 
                                      ssp. brachyanthum
                                                             dwarf
                                      ssp. hypolepidotum
                       R. charitopes
                                      ssp. charitopes
                                      ssp. tsangpoense
                                              var. curvistylum
                       R. glaucophyllum
                                              var. album
                                              var. glaucophyllum
                                              var. tubiforme  -> R.tubiforme*
                       R. luteiflorum
                       R. pruniflorum
                       R. shweliense
               
               Subsection Heliolepida
 
                       R. bracteatum
                       R. heliolepis
                                              var. brevistylum
                                              var. fumidum
                                              var. heliolepis 
                                              var. oporinum*         
                       R. hirsutipetiolatum*
                       R. invictum*
                       R. rubiginosum 
                                              var. rubiginosum*
                                              var. ptilostylum
                                                             desquamatum
                       
               Subsection Lapponica
 
                       R. amundsenianum
                       R. bamaense*
                       R. bulu
                       R. burjaticum
                       R. capitatum
                       R. complexum                                  
                       R. cuneatum
                       R. dasypetalum
                       R. dawuense*
                       R. declivatum*
                      R. x edgarianum
                       R. fastigiatum
                       R. flavidum
                       R. gologense*
                       R. hippophaeoides
                                              var. hippophaeoides
                                              var. occidentale (fimbriatum)
                       R. impeditum
                                                             litangense
                       R. intricatum
                       R. joniense*
                       R. labolengense*
                       R. lapponicum
                                                             parvifolium
                       R. lungchiense*
                       R. maowenense*
                       (R. ninyyaense)
                       R. nitidulum
                                              var. nitidulum
                                              var. omeiense
                       R. nivale
                                      ssp. australe
                                      ssp. boreale           f.  nigropunctatum, stictophyllum
                                                                 BeimaShan, 
                                      ssp. nivale
                       R. orthocladum
                                              var. microleucum       
                                              var. orthocladum
                                              var. longistylum*
        
                       R. polycladum                                 f. scintillans
                       R. qinghaiense*
                       R. rupicola
                                              var. rupicola
                                              var. chryseum
                                              var. muliense
                       R. russatum
                       R. setosum
                       R. taibaiense*
                       R. tapetiforme
                       R. telmateium
                       R. thymifolium
                       R. tsaii
                       (R x verruculosum)
                       R. websterianum
                                              var. websterianum
                                              var. yulongense
                       R. xiguense*
                       R. yulingense
                       R. yungningense
                       R. yushuense*
                       R. zekoense*
                       R. zheguense*
                       
 
               Subsection Ledum*  http://www.rhododendron.dk/Ledum.html
 
                       R. diversipilosum              
                       R. groenlandicum
                       R. hypoleucum
                       R. neoglandulosum
                       R. subarcticum 
                       R. subulatum
                       R. tolmachevii
                       R. tomentosum  
        
               Subsection Lepidota
 
                       R. cowanianum
                       R. lepidotum
                       R. lowndesii
 
               Subsection Maddenia
 
                 Maddenii alliance
                        R. maddenii
                                      ssp. crassum
                                                             chapaense
                                                             subranubium
                                       ssp. maddenii
                 Dalhousiae alliance  
                       R. chunienii*
                       R. dalhousiae          
                                              var. dalhousiae
                                              var. rhabdotum
                       R. excellens
                       R. headfortianum
                       R. kiangsiense *
                       R. levinei
                       R. liliiflorum
                       R. lindleyi
                       R. nuttallii
                       R. sinonuttallii*
                       R. taggianum
                 Megacalyx alliance   
                       R. megacalyx
                 Ciliicalyx / Johnstoneanum alliance
                       R. changii*
                       R. burmanicum
                       R. ciliatum
                       R. coxianum
                       R. fletcherianum
                       R. formosum            var. formosum
                                                                     iteophyllum
                                              var. inaequale
                       R. scopulorum
                       R. valentinianum
                                              var. changii
                                              var. oblongilobatum*
                                              var. valentinianum*
                       R. yungshangense*
                 Johnstoneanum aggregate      
                       R. dendricola
                       R. johnstoneanum
                       R. walongense
                 Ciliicalyx aggregate 
                       R. carneum
                       R. ciliicalyx*
                                              ssp. ciliicalyx*
                                              ssp. lyi*
                       R. horlickianum
                       R. leptocladon*
                       R. ludwigianum
                       R. pachypodum
                       R. parryae
                       R. roseatum
                       R. veitchianum
                                                     cubittii
                       R. pseudociliipes
                       -----
                       R. linearilobum*
                       R. zihangense*
                       R. yaogangxianense*
                       R. ciliipes*
                       R. pseudociilipes*
                       R. rhombifolium*
                       R. taronense*
                       R. wumingense*
                       R. scopulorum*
                       -----
                       R. amandum
                       R. crenulatum
                       R. cuffeanum
                       R. fleuryi
                       R. goreri
                       R. grothausii
                       R. mianningense (Lapponica)
                       R. nemorosum
                       R. rufosquamosum
                       R. surasianum
                       R. yungchangense
 
               Subsection Micrantha 
                       R. brevicaudatum*
                       R. liaoxigense*
                       R. micranthum
 
               Subsection Monantha*
                       R. concinnoides
                       R. flavantherum
                       R. kasoense*
                       R. monanthum*
 
               Subsection Moupinensia
 
                       R. dendrocharis
                       R. moupinense
                       R. petrocharis*
                       
                Subsection Rhododendron
 
                       R. ferrugineum
                       R. x intermedium ?
                       R. hirsutum
                       R. myrtifolium
 
               Subsection Rhodorastra
 
                       R. dauricum
                       R. ledebourii
                       R. mucronulatum 
                                      var. mucronulatum
                                      var. taquetii
                       R. sichotense
 
               Subsection Saluenensia
 
                       R. calostrotum 
                               ssp. calostrotum
                                      var. calostrotum*
                                      var. calciphilum*
                               ssp. keleticum             radicans 
                               ssp. riparioides
                               ssp. riparium              nitens
                                                     
                       R. saluenense                  
                               ssp. chameunum             prostratum
                               ssp. saluenense        
 
               Subsection Scabrifolia  (à subgenus Pseudorhodorastrum*
                                              sec. Rhabdorhodion*
                                              sec. Rhodobotrys*)
                       R. hemitrichotum
                       R. mollicomum
                       R. pubescens
                       R. racemosum   à sec. Rhodobotrys*
                       R. fuyuanense* à sec. Rhodobotrys*
                       R. scabrifolium 
                                      var. pauciflorum
                                      var. scabrifolium
                                      var. spiciferum  àR. speciferum*
                       R. spinuliferum
                                      var. glabrescens*
                                      var. spiciferum  àR. speciferum *
 
               Subsection Tephropepla
 
                       R. auritum
                       R. hanceanum
                       R. longistylum
                       R. tephropeplum
                       R. tsinlingense*
                       R. xanthostephanum
 
               Subsection Trichoclada  à Pseudazalea*
 
                       R. caesium
                       R. lepidostylum
                       R. mekongense
                                      var. longipilosum*
                                      var. mekongense
                                      var. melinanthum*
                                      var. rubrolineatum
                       R. nanjianense*
                       R. trichocladum
                       R. viridescens
                                                      rubroluteum
               Subsection Triflora
                       
                  Augustinii / Triflorum alliance
                       R. ambiguum    
                                                     chengshienanum
                       R. amesiae
                       R. augustinii
                               ssp. augustinii
                               ssp. chasmanthum
                               ssp. hardyi
                               ssp. rubrum
 
                       R. bivelatum
                       R. brachypodum*
                       R. concinnum   
                                      var. benthamianum
                                                     pseudoyanthinum
                               (ssp. gonggaense in limbo H.E.)
                       R. gemmiferum*
                       R. guangnanense*
                       R. keiskei
                                      var. hypoglaucum
                                      var. keiskei
                                      var. ozawae
                       R. lutescens
                       R. polylepis
                       R. searsiae
                       R. seguinii
                       R. shaanxiense
                       R. shimianense*
                       R. trichanthum
                       R. triflorum
                                      var. bauhiniiflorum  (ssp. multiflorum*)
                                      var. triflorum       (ssp. triflorum*)
                       R. wongii*
                       R. zaleucum
                                      var. flaviflorum
                                      var. pubiflorum*
                                      var. zaleucum
                  Yunnanense alliance 
                       R. davidsonianum
                       R. pleistanthum*
                       R. rigidum
                       R. siderophyllum
                       R. tatsienense
                                      var. tatseriense*  (R. kangdingense)
                                      var. nudatum*
                       R. xichangense*
                       R. yunnanense
                                                     bodinieri 
                                                     subruseum 
                                                     hormophorum 
                       
                  Oreotrephes alliance        
                       R. oreotrephes
 
                  Subsection Uniflora
                       R. imperator
                       R. ludlowii
                       R. pemakoense
                       R. pumilum
                       R. uniflorum
 
                  Subsection Virgata  (monotypic)  àRhabdorhodion*
 
                       R. virgatum    
                               ssp. virgatum
                               ssp. oleifolium
 
        Section Schistanthe (vireya)        Section 'vireya' not updated look at   press for species
Only two subsection found with DNA analysis: Pseudovireya and Euvireya (all the other groups)   !Phylogenetic-DNA tree
 
Subsection Albovireya
 
R. aequabile
R. album
R. arenicolum
R. cernuum
R. comptum
                var. trichodes
R. correoides
R. giulianetti
R. lagunculicarpum
R. lampongum
R. proliferum
R. pudorinum
R. versteegii
R. yelliotii
R. zollingeri
 
Subsection Malayovireya
 
R. acuminatum
R. apoanum
R. durionifolium
         ssp. sabahense
R. fallacinum
R. fortunans
R. himantodes
                var. lavandulifolium
R. lamrialianum
                var. gunsalamianum
R. lineare
R. malayanum
                var. axillare
                var. axillare latifolium
                var. axillare ovatum
                var. infrapilosum
                var. pilosifilum
                var. pubens
R. micromalayanum
R. nortoniae
R. obscurum
R. variolosum
                var. andersonii
R. vinicolor
 
 
Subsection Phaeovireya
 

R.asperrimum

 R. asperum

R.beyerinckianum

R.bryophilum

R.bullifolium

R.caliginis

R.cyatheicolum

R.delicatulum

                var.lanceolatoides

R.dianthosmum

R.dielsianum

                var.stylotrichum

R.extrorsum

R.eymae

R.gardenia

R.gilliardii

R.haematophthalmum

R.hellwigii

R.hooglandii

R.hyacinthosmum

R.kerowagiense

                                      var.phaeopeplum

R.leptanthum

                                      var.warianum

R.melantherum

R.neriifolium

R.opulentum

R.phaeochitum

R.phaeochristum

R.phaeops

R.prainianum

R.psilanthum

R.rappardii

R.rarum

R.revolutum

R.rhodochroum

R.rubellum

R.schoddei

R.solitarium

R.spondylophyllum

R.stelligerum

R.stolleanum

R.superbum

R.thaumasianthum

R.truncicolum

R.tuberculiferum


Subsection Discovireya
 

R.asperrimum

R.asperum

R.beyerinckianum

R.bryophilum

R.bullifolium

R.caliginis

R.cyatheicolum

R.delicatulum

                                      var.lanceolatoides

R.dianthosmum

R.dielsianum

                                      var.stylotrichum

R.extrorsum

R.eymae

R.gardenia

R.gilliardii

R.haematophthalmum

R.hellwigii

R.hooglandii

R.hyacinthosmum

R.kerowagiense

                                      var.phaeopeplum

R.leptanthum

                                      var.warianum

R.melantherum

R.neriifolium

R.opulentum

R.phaeochitum

R.phaeochristum

R.phaeops

R.prainianum

R.psilanthum

R.rappardii

R.rarum

R.revolutum

R.rhodochroum

R.rubellum

R.schoddei

R.solitarium

R.spondylophyllum

R.stelligerum

R.stolleanum

R.superbum

R.thaumasianthum

R.truncicolum

R.tuberculiferum

SubsectionPseudovireya

 

 

R.adinophyllum

R.asperulum

R.borneense

                               ssp.angustissimum

                               ssp.villosum

R.buxoides

R.capellae

R.ciliilobum

R.cuneifolium

                                              var.microcarpum

R.cyrtophyllum

R.datiandingense

R.densifolium

R.emarginatum

                                              var.eriocarpum

R.ericoides

R.erosipetalum

R.gaultheriifolium

                                              var.expositum

R.hameliiflorum

R.insculptum

R.kawakamii

R.leiboense

R.lindaueanum

                                              var.bantaengense

R.maguanense

R.meliphagidum

R.nanophyton

                                              var.petrophilum

R.nummatum

R.oreites

                                              var.chlorops

R.perakense

R.pulleanum

                                              var.maiusculum

R.quadrasianum

                                              var.davaoense

                                              var.intermedium

                                              var.malindangense

                                              var.marivelesense

                                              var.rosmarinifolium

                                              var.selebicum

R.retusum

                                              var.epilosum

                                              var.trichostylum

R.rupivalleculatum

R.rushforthii

R.santapaui

R.saruwagedicum

R.schizostigma

R.scortechinii

R.seimundii

R.sororium

R.spathulatum

R.taxoides

R.vaccinioides

R.vanderbiltianum

R.vinkii

 

Subsection Siphonovireya

 

 

R.agathodaemonis

R.cinchoniflorum

R.gideonii

R.habbemae

R.herzogii

R.incommodum

R.inundatum

R.protandrum

R.reevei

 

Subsection Solenovireya

 

R.alborugosum

R.amabile

R.archboldianum

R.armitii

R.brachypodarium

R.carrii

R.carringtoniae

R.carstensense

R.chamaepitys

R.cinerascens

R.cruttwellii

R.edanoi

                               ssp.pneumonanthum

R.goodenoughii

R.hartleyi

R.jasminiflorum

                                              var.copelandii

                                              var.heusseri

                                              var.oblongifolium

                                              var.punctatum

R.lambianum

R.loranthiflorum

                               ssp.lakekamuensis

R.macrosiphon

R.maius

R.mendumiae

R.mogeanum

R.multinervium

R.natalicium

R.niveoflorum

R.oliganthum

R.orbiculatum

R.oreadum

R.pleianthum

R.pseudotricanthum

R.pubitubum

R.radians

                                              var.minahasae

R.retrorsipilum

R.rhodoleucum

R.rhodosalpinx

R.roseiflorum

R.ruttenii

R.searleanum

R.stapfianum

R.suaveolens

                                              var.roseum

R.subpacificum

R.syringoideum

R.tuba

 

Subsection Vireya

 

R.abietifolium

R.acrophilum

R.alternans

R.alticolum

R.anagalliflorum

R.angulatum

R.apiense

R.arfakianum

R.atropurpureum

R.aurigeranum

                               ssp.hirsutum

R.baconii

R.baenitzianum

R.bagobonum

R.banghamiorum

R.beccarii

R.blackii

R.bloembergenii

R.brachygynum

R.brassii

R.brevipes

R.burttii

R.buruense

R.buxifolium

R.caespitosum

R.calosanthes

R.celebicum

R.chevalieri

R.christi

R.christianae

R.citrinum

                                              var.discoloratum

R.coelorum

R.commonae

R.commutatum

R.comparabile

R.cornu-bovis

R.crassifolium

R.culminicolum

                                              var.angiense

                                              var.nubicola

R.curviflorum

R.cuspidellum

R.detznerianum

R.disterigmoides

                               ssp.astromontium

R.englerianum

R.exuberans

R.flavoviride

R.frey-wysslingii

R.glabriflorum

R.gracilentum

R.hatamense

R.helodes

R.hirtolepidotum

R.impositum

R.impressopunctatum

R.inconspicuum

R.intranervatum

R.javanicum

                                ssp.brookeanum

                                ssp.cockburnii

                                ssp.palawanense

                                ssp.schadenbergii

                                              var.teysmannii

R.kemulense

R.kochii

R.laetum

R.lamii

R.lanceolatum

R.leptobrachion

R.leptomorphum

R.leucogigas

R.leytense

                                              var.loheri

R.loboense

R.lochiae

R.loerzingii

R.lompohense

R.longiflorum

                                              var.bancanum

                                              var.longipetalum

                                              var.subcordatum

R.lowii

R.luraluense

R.luteosquamatum

R.macgregoriae

                        var.glabrifilum

                                              var.mayrii

R.madulidii

R.maxwellii

R.meijeri

R.microphyllum

R.mindanaense

R.mollianum

R.monkoboense

R.multicolor

R.muscicola

R.myrsinites

R.neobritannicum

R.nervulosum

R.nieuwenhuisii

R.oxycoccoides

R.pachycarpon

R.pachystigma

R.papuanum

R.parvulum

R.pauciflorum

                                              var.calocodon

R.perplexum

R.polyanthemum

R.poremense

R.porphyranthes

R.praetervisum

R.psammogenes

R.pseudobuxifolium

R.pseudomurudense

R.pubigermen

R.purpureiflorum

R.pusillum

R.pyrrhophorum

R.rarilepidotum

                                              var.ootrichum

R.renschianum

R.retivenium

R.reynosoi

R.rhodopus

R.rhodostomum

R.ripleyi

                                              var.basitrichum

                                              var.cryptogonium

R.robinsonii

R.rosendahlii

R.rousei

R.rubineiflorum

R.rubrobracteatum

R.rugosum

                                              var.kinabaluense

R.salicifolium

R.sarcodes

R.saxifragoides

R.sayeri

R.scabridibracteum

R.scarlatinum

R.schlechteri

R.seranicum

R.sessilifolium

R.simulans

R.stenophyllum

                              ssp.angustifolium

R.stevensianum

R.stresemanii

R.subcrenulatum

R.subuliferum

R.subulosum

R.sumatranum

R.takeuchii

R.taxifolium

R.toxopei

R.triumphans

R.tuhanensis

R.ultimum

R.vanvuurenii

R.verticillatum

                                              var.velutinum

R.vidalii

                              ssp.brachystemon

R.villosulum

R.viriosum

R.vitis-idaea

R.wentianum

R.whiteheadii

R.wilkiei

R.williamsii

R.womersleyi

R.wrightianum

                                              var.cyclopense

                                              var.insulare

R.xanthopetalum

R.xenium

R.yongii

R.zoelleri

 
 
 
               
                

Subgenus Therorhodion

(probably not in this genus H.Eiberg 1998)
 
                       R. camtschaticum
                               ssp. camtschaticum
                               ssp. glandulosum
                       R. redowskianum
 
        
 
-------------------------------------------------------------------------------------------------
Title: Taxonomic revision of Rhododendron mariae (Ericaceae) and related taxa

 
 
 
Authors: Jin, XF; Ding, BY; Jin, SH; Qian, L

 
 
 
Author Full Names: Jin, Xiao-Feng; Ding, Bing-Yang; Jin, Shui-Hu; Qian, Li

 
 
 
Source: NORDIC JOURNAL OF BOTANY, 27 (3): 186-202 2009

 
 
 
Language: English

 
 
 
Document Type: Article

 
 
 
Abstract: 

 
 
 
Based on a literature survey and a field trip with specimen examination, statistical analysis including principal components analysis (PCA) on morphological characters have been carried out on Rhododendron mariae and related taxa. Cluster analysis and PCA showed three independent groups, which corresponded to three species except R. loniceriflorum. As a consequence, nine taxa are reduced to synonyms of R. mariae, and two new combinations, viz R. mariae subsp. microphyton, and R. mariae subsp. flumineum are made. Rhododendron microphyton var. trichanthum is reduced to a synonym of R. mariae subsp. microphyton, and R. malipoense to a synonym of R. mariae subsp. flumineum. Three names are synonymized and a new combination, viz R. chryocalyx subsp. meridionale, is proposed within R. chryocalyx. The third entity, R. naamkwanense, has got two new synonyms. Rhododendron loniceriflorum, with whitish pubescence on the inner corolla surface, is recognized as the fourth distinct spe! cies. Each entity is illustrated with line drawings for identification